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Heterodermia pseudospeciosa (Kurok.) Culb.

Family :


W. Culberson, Bryologist 69: 484. 1966 

Basionym: Anaptychia pseudospeciosa Kurokawa, l. lap. Bot. 34: 176. 1959

ID :


Thallus :

Foliose, orbicular to irregularly spreading, adnate to loosely adnate, 2–5 cm wide

Lobes :

Comparatively small, 0.7–1.5 mm wide, plane to weakly convex, sublinear-elongate, sympodial to subdigitately branched, usually discrete at the periphery, radiating; apices not ascending, eciliate, with distinct pseudocyphellae along the margins of the lobes or on small short lateral branches

Upper Side:

Grayish white to gray to brownish gray, ± darker at the lobe tips, pruinose or not

Isidia :

Sorediate; soralia white to bluish gray, originating mostly from bursting pseudocyphellae, developing into subcapitate or labriform soralia.

Lower Side :

Corticate, whitish to pale brown, rarely dark gray in the center. Lower surface rhizines sparse, mostly simple or sparsely branched, marginal rhizines frequent, concolorous with the thallus or becoming dark or black near the apices, irregularly branched, to 1 mm long, projecting beyond the lobes

Medulla :


Rare, laminal, adnate to sessile, 1–3 mm wide; margin initially crenulate, later sorediate; disc concave, brown to brown-black, epruinose

Apothecia :

Pachysporaria-type, ellipsoidal, 26–32 × 12–14 μm. Pycnidia immersed or slightly protruding; conidia bacilliform, 4–5 × 1 μm

Ascospores :

Cortex K+ yellow, C–, KC–, P+ yellow; medulla K+ yellow then red, C–, P+ dark yellow; containing atranorin (major), zeorin (major), 16β-acetoxyhopane-6α,22-diol (major), leucotylin (minor), 6αacetoxyhopane-16β,22-diol (trace), norstictic acid (major), connorstictic acid (minor), salazinic acid (trace)

Chemistry :

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